Urocystis violae (Sowerby) A.A. Fisch. Waldh.
(Violet Smut)

Interactions where Urocystis violae is the dominant partner and gains from the process

The following relationships have been collated from the published literature (see 'Interaction References').

Stage		Summary			Taxon	Vernacular	Classification	References	Darwin Classification of Active Taxon	Active Taxon	Active Vernacular	Active Taxon Uncertain	Active State	Active Part	Active Stage	Relationship	Relationship Uncertain	Relationship Geography	Darwin Classification of Passive Taxon	Passive Taxon	Passive Vernacular	Passive Taxon Uncertain	Passive State	Passive Part	Passive Stage	Passive Taxon's significance to Active Taxon	Indoors etc	Season
spore mass		spore mass causes gall of live petiole			Viola hirta	Hairy Violet	Malpighiales: Violaceae	Checklist of the British and Irish Basidiomycota, Legon, N.W. & Henrici, A. with Roberts, P.J., Spooner, B.M. & Watling, R., 2005	/Fungi/Basidiomycota/Ustilaginomycetes/Urocystidales/Urocystidaceae/Urocystis violae/	Urocystis violae	violet smut				spore mass	Foodplant / gall		UK and/or Eire	/Plantae/Tracheophyta/Magnoliopsida/Malpighiales/Violaceae/Viola hirta/	Viola hirta	hairy violet		live	petiole				(11-7)
spore mass		spore mass causes gall of live leaf vein			Viola hirta	Hairy Violet	Malpighiales: Violaceae	Checklist of the British and Irish Basidiomycota, Legon, N.W. & Henrici, A. with Roberts, P.J., Spooner, B.M. & Watling, R., 2005	/Fungi/Basidiomycota/Ustilaginomycetes/Urocystidales/Urocystidaceae/Urocystis violae/	Urocystis violae	violet smut				spore mass	Foodplant / gall		UK and/or Eire	/Plantae/Tracheophyta/Magnoliopsida/Malpighiales/Violaceae/Viola hirta/	Viola hirta	hairy violet		live	leaf vein				(11-7)
spore mass		spore mass causes gall of live rootstock (upper parts)			Viola hirta	Hairy Violet	Malpighiales: Violaceae	Checklist of the British and Irish Basidiomycota, Legon, N.W. & Henrici, A. with Roberts, P.J., Spooner, B.M. & Watling, R., 2005	/Fungi/Basidiomycota/Ustilaginomycetes/Urocystidales/Urocystidaceae/Urocystis violae/	Urocystis violae	violet smut				spore mass	Foodplant / gall		UK and/or Eire	/Plantae/Tracheophyta/Magnoliopsida/Malpighiales/Violaceae/Viola hirta/	Viola hirta	hairy violet		live	rootstock (upper parts)				(11-7)
spore mass		spore mass causes gall of live petiole			Viola odorata	Sweet Violet	Malpighiales: Violaceae	Microfungi on Land Plants: An Identification Handbook, Ellis, M.B. & J.P., 1997 Checklist of the British and Irish Basidiomycota, Legon, N.W. & Henrici, A. with Roberts, P.J., Spooner, B.M. & Watling, R., 2005 Provisional Keys to British Plant Galls, Stubbs, F.B. (Editor), 1986	/Fungi/Basidiomycota/Ustilaginomycetes/Urocystidales/Urocystidaceae/Urocystis violae/	Urocystis violae	violet smut				spore mass	Foodplant / gall		UK and/or Eire	/Plantae/Tracheophyta/Magnoliopsida/Malpighiales/Violaceae/Viola odorata/	Viola odorata	sweet violet		live	petiole				(11-7)
spore mass		spore mass causes gall of live rootstock (upper parts)			Viola odorata	Sweet Violet	Malpighiales: Violaceae	Microfungi on Land Plants: An Identification Handbook, Ellis, M.B. & J.P., 1997 Checklist of the British and Irish Basidiomycota, Legon, N.W. & Henrici, A. with Roberts, P.J., Spooner, B.M. & Watling, R., 2005 Provisional Keys to British Plant Galls, Stubbs, F.B. (Editor), 1986	/Fungi/Basidiomycota/Ustilaginomycetes/Urocystidales/Urocystidaceae/Urocystis violae/	Urocystis violae	violet smut				spore mass	Foodplant / gall		UK and/or Eire	/Plantae/Tracheophyta/Magnoliopsida/Malpighiales/Violaceae/Viola odorata/	Viola odorata	sweet violet		live	rootstock (upper parts)				(11-7)
spore mass		spore mass causes gall of live leaf vein			Viola odorata	Sweet Violet	Malpighiales: Violaceae	Microfungi on Land Plants: An Identification Handbook, Ellis, M.B. & J.P., 1997 Checklist of the British and Irish Basidiomycota, Legon, N.W. & Henrici, A. with Roberts, P.J., Spooner, B.M. & Watling, R., 2005 Provisional Keys to British Plant Galls, Stubbs, F.B. (Editor), 1986	/Fungi/Basidiomycota/Ustilaginomycetes/Urocystidales/Urocystidaceae/Urocystis violae/	Urocystis violae	violet smut				spore mass	Foodplant / gall		UK and/or Eire	/Plantae/Tracheophyta/Magnoliopsida/Malpighiales/Violaceae/Viola odorata/	Viola odorata	sweet violet		live	leaf vein				(11-7)
spore mass		spore mass causes gall of live petiole			Viola reichenbachiana	Early Dog-violet	Malpighiales: Violaceae	Microfungi on Land Plants: An Identification Handbook, Ellis, M.B. & J.P., 1997 Checklist of the British and Irish Basidiomycota, Legon, N.W. & Henrici, A. with Roberts, P.J., Spooner, B.M. & Watling, R., 2005 Provisional Keys to British Plant Galls, Stubbs, F.B. (Editor), 1986	/Fungi/Basidiomycota/Ustilaginomycetes/Urocystidales/Urocystidaceae/Urocystis violae/	Urocystis violae	violet smut				spore mass	Foodplant / gall		UK and/or Eire	/Plantae/Tracheophyta/Magnoliopsida/Malpighiales/Violaceae/Viola reichenbachiana/	Viola reichenbachiana	early dog-violet		live	petiole				(11-7)
spore mass		spore mass causes gall of live rootstock (upper parts)			Viola reichenbachiana	Early Dog-violet	Malpighiales: Violaceae	Microfungi on Land Plants: An Identification Handbook, Ellis, M.B. & J.P., 1997 Checklist of the British and Irish Basidiomycota, Legon, N.W. & Henrici, A. with Roberts, P.J., Spooner, B.M. & Watling, R., 2005 Provisional Keys to British Plant Galls, Stubbs, F.B. (Editor), 1986	/Fungi/Basidiomycota/Ustilaginomycetes/Urocystidales/Urocystidaceae/Urocystis violae/	Urocystis violae	violet smut				spore mass	Foodplant / gall		UK and/or Eire	/Plantae/Tracheophyta/Magnoliopsida/Malpighiales/Violaceae/Viola reichenbachiana/	Viola reichenbachiana	early dog-violet		live	rootstock (upper parts)				(11-7)
spore mass		spore mass causes gall of live leaf vein			Viola reichenbachiana	Early Dog-violet	Malpighiales: Violaceae	Microfungi on Land Plants: An Identification Handbook, Ellis, M.B. & J.P., 1997 Checklist of the British and Irish Basidiomycota, Legon, N.W. & Henrici, A. with Roberts, P.J., Spooner, B.M. & Watling, R., 2005 Provisional Keys to British Plant Galls, Stubbs, F.B. (Editor), 1986	/Fungi/Basidiomycota/Ustilaginomycetes/Urocystidales/Urocystidaceae/Urocystis violae/	Urocystis violae	violet smut				spore mass	Foodplant / gall		UK and/or Eire	/Plantae/Tracheophyta/Magnoliopsida/Malpighiales/Violaceae/Viola reichenbachiana/	Viola reichenbachiana	early dog-violet		live	leaf vein				(11-7)
spore mass		spore mass causes gall of live rootstock (upper parts)			Viola riviniana	Common Dog-violet, Blunt-spurred Violet	Malpighiales: Violaceae	Microfungi on Land Plants: An Identification Handbook, Ellis, M.B. & J.P., 1997 Checklist of the British and Irish Basidiomycota, Legon, N.W. & Henrici, A. with Roberts, P.J., Spooner, B.M. & Watling, R., 2005 Provisional Keys to British Plant Galls, Stubbs, F.B. (Editor), 1986	/Fungi/Basidiomycota/Ustilaginomycetes/Urocystidales/Urocystidaceae/Urocystis violae/	Urocystis violae	violet smut				spore mass	Foodplant / gall		UK and/or Eire	/Plantae/Tracheophyta/Magnoliopsida/Malpighiales/Violaceae/Viola riviniana/	Viola riviniana	common dog-violet, blunt-spurred violet		live	rootstock (upper parts)				(11-7)
spore mass		spore mass causes gall of live leaf vein			Viola riviniana	Common Dog-violet, Blunt-spurred Violet	Malpighiales: Violaceae	Microfungi on Land Plants: An Identification Handbook, Ellis, M.B. & J.P., 1997 Checklist of the British and Irish Basidiomycota, Legon, N.W. & Henrici, A. with Roberts, P.J., Spooner, B.M. & Watling, R., 2005 Provisional Keys to British Plant Galls, Stubbs, F.B. (Editor), 1986	/Fungi/Basidiomycota/Ustilaginomycetes/Urocystidales/Urocystidaceae/Urocystis violae/	Urocystis violae	violet smut				spore mass	Foodplant / gall		UK and/or Eire	/Plantae/Tracheophyta/Magnoliopsida/Malpighiales/Violaceae/Viola riviniana/	Viola riviniana	common dog-violet, blunt-spurred violet		live	leaf vein				(11-7)
spore mass		spore mass causes gall of live petiole			Viola riviniana	Common Dog-violet, Blunt-spurred Violet	Malpighiales: Violaceae	Microfungi on Land Plants: An Identification Handbook, Ellis, M.B. & J.P., 1997 Checklist of the British and Irish Basidiomycota, Legon, N.W. & Henrici, A. with Roberts, P.J., Spooner, B.M. & Watling, R., 2005 Provisional Keys to British Plant Galls, Stubbs, F.B. (Editor), 1986	/Fungi/Basidiomycota/Ustilaginomycetes/Urocystidales/Urocystidaceae/Urocystis violae/	Urocystis violae	violet smut				spore mass	Foodplant / gall		UK and/or Eire	/Plantae/Tracheophyta/Magnoliopsida/Malpighiales/Violaceae/Viola riviniana/	Viola riviniana	common dog-violet, blunt-spurred violet		live	petiole				(11-7)
spore mass		spore mass causes gall of live petiole			[Viola sp. cult.]	cultivated violets	Malpighiales: Violaceae	Microfungi on Land Plants: An Identification Handbook, Ellis, M.B. & J.P., 1997 Checklist of the British and Irish Basidiomycota, Legon, N.W. & Henrici, A. with Roberts, P.J., Spooner, B.M. & Watling, R., 2005 Provisional Keys to British Plant Galls, Stubbs, F.B. (Editor), 1986	/Fungi/Basidiomycota/Ustilaginomycetes/Urocystidales/Urocystidaceae/Urocystis violae/	Urocystis violae	violet smut				spore mass	Foodplant / gall		UK and/or Eire	/Plantae/Tracheophyta/Magnoliopsida/Malpighiales/Violaceae/Viola sp. cult./	Viola sp. cult.	cultivated violets		live	petiole				(11-7)
spore mass		spore mass causes gall of live rootstock (upper parts)			[Viola sp. cult.]	cultivated violets	Malpighiales: Violaceae	Microfungi on Land Plants: An Identification Handbook, Ellis, M.B. & J.P., 1997 Checklist of the British and Irish Basidiomycota, Legon, N.W. & Henrici, A. with Roberts, P.J., Spooner, B.M. & Watling, R., 2005 Provisional Keys to British Plant Galls, Stubbs, F.B. (Editor), 1986	/Fungi/Basidiomycota/Ustilaginomycetes/Urocystidales/Urocystidaceae/Urocystis violae/	Urocystis violae	violet smut				spore mass	Foodplant / gall		UK and/or Eire	/Plantae/Tracheophyta/Magnoliopsida/Malpighiales/Violaceae/Viola sp. cult./	Viola sp. cult.	cultivated violets		live	rootstock (upper parts)				(11-7)
spore mass		spore mass causes gall of live leaf vein			[Viola sp. cult.]	cultivated violets	Malpighiales: Violaceae	Microfungi on Land Plants: An Identification Handbook, Ellis, M.B. & J.P., 1997 Checklist of the British and Irish Basidiomycota, Legon, N.W. & Henrici, A. with Roberts, P.J., Spooner, B.M. & Watling, R., 2005 Provisional Keys to British Plant Galls, Stubbs, F.B. (Editor), 1986	/Fungi/Basidiomycota/Ustilaginomycetes/Urocystidales/Urocystidaceae/Urocystis violae/	Urocystis violae	violet smut				spore mass	Foodplant / gall		UK and/or Eire	/Plantae/Tracheophyta/Magnoliopsida/Malpighiales/Violaceae/Viola sp. cult./	Viola sp. cult.	cultivated violets		live	leaf vein				(11-7)

Author & Year	Title	Source
Ellis, M.B. & J.P., 1997	Microfungi on Land Plants: An Identification Handbook	2nd (New Enlarged) edition, 868pp, The Richmond Publishing Co. Ltd
Legon, N.W. & Henrici, A. with Roberts, P.J., Spooner, B.M. & Watling, R., 2005	Checklist of the British and Irish Basidiomycota	517pp, Royal Botanic Gardens, Kew
Stubbs, F.B. (Editor), 1986	Provisional Keys to British Plant Galls	95pp, British Plant Gall Society (BPGS)

Urocystis violae (Violet Smut) may also be included in 'feeds on' relations listed under the following higher taxa:

FUNGI
(true fungi)

Urocystis violae (Violet Smut) may be included in 'fed on by' relations listed under the following higher taxa:

FUNGI
(true fungi)

Fungi s.l.
(fungi (in the broad sense))

Taxonomic hierarchy:

Species	Urocystis violae (Violet Smut)
Genus	Urocystis (a genus of smut fungi)
Family	UROCYSTIDACEAE (a family of fungi)
Order	UROCYSTIDALES (an order of fungi)
Class	USTILAGINOMYCETES (smut fungi and relatives)
Subphylum	USTILAGINOMYCOTINA (a subphylum of fungi)
Phylum	BASIDIOMYCOTA (spore droppers, basidiomycetes)
Kingdom	FUNGI (true fungi)
Domain	Eukaryota (eukaryotes)
Life	BIOTA (living things)

Urocystis violae is also a member of:
Micromycetes - microfungi (via Ustilaginomycetes)
Fungi s.l. - fungi (in the broad sense) (via Fungi)
Lower Plants - lower plants (via Fungi s.l.)

NBN

NBN (data.nbn.org.uk) has a distribution map for Urocystis violae (Violet Smut)

Identification Works

Handling & Magnification	Author	Year	Title	Source
	Mordue, J.E.M.	1970	Urocystis violae	I.M.I. Descriptions of Pathogenic Fungi and Bacteria CMI Descriptions of Pathenogenic Fungi and Bacteria Sheet 969.

Urocystis violae (Violet Smut) may also be included in identification literature listed under the following higher taxa:

USTILAGINOMYCETES
(smut fungi and relatives)

USTILAGINOMYCOTINA
(a subphylum of fungi)

BASIDIOMYCOTA
(spore droppers, basidiomycetes)

FUNGI
(true fungi)

Eukaryota
(eukaryotes)

BIOTA
(living things)

Fungi s.l.
(fungi (in the broad sense))

Literature listed under the following higher taxa may be relevant to Urocystis violae (Violet Smut):

USTILAGINOMYCETES
(smut fungi and relatives)

USTILAGINOMYCOTINA
(a subphylum of fungi)

BASIDIOMYCOTA
(spore droppers, basidiomycetes)

FUNGI
(true fungi)

Eukaryota
(eukaryotes)

BIOTA
(living things)

Fungi s.l.
(fungi (in the broad sense))

Widespread, many records.

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